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Rowan Lockwood

Professor of Geology

Office: McGlothlin-Street Hall 226
Email: [[rxlock]]
Office Phone: 757 221 2878
Research Areas: Paleontology, Paleobiology, Cenozoic Global Climate Change, Evolution/Ecology of Fossil Marine Invertebrates

  • Ph.D., University of Chicago
  • M.Sc., University of Bristol, UK
  • B.A., Yale University
Research Interests

The overall goal of my research is to understand how extinction and environmental change influence the evolution and ecology of fossil marine invertebrates. My research interests are broad and range from the influence of the K/T mass extinction on bivalve evolution, to the taphonomy and historical ecology of Holocene benthic mollusks in the Chesapeake Bay, to the effects of early Cenozoic global climate change during the early Cenozoic on venericard bivalves along the Gulf Coastal Plain.

Current Projects
  • Conservation paleobiology of Chesapeake Bay oysters: using the fossil and archaeological records to establish baselines for restoration.
  • Extinction risk in ancient seas: using the fossil record to predict the response of marine organisms to climate change and ocean acidification.
  • Evolutionary and ecological response of venericard bivalves to climate change during the Paleogene.
  • Application of the Paleobiology Database to teaching paleontology at the undergraduate level.
Selected Publications

* denotes undergraduate author; † denotes graduate student author; ‡ denotes post-doctoral fellow

*Zimmt, J. B., S. M. Kidwell, R. Lockwood, and M. Thirlwall. 2022. Strontium isotope stratigraphy reveals 100 ky-scale condensation, beveling, and internal shingling of transgressive shell beds in the Maryland Miocene, Palaios, 37:553–573. Download pdf Copy.

*Chriscoe, M., R. Lockwood, J. S. Tweet, and V. L. Santucci. 2022. Colonial National Historical Park Paleontological Inventory. Natural Resource Report NPS/COLO/NRR—2022/2361. Download pdf Copy.

*Samuels-Fair, M., M. João Fernandes Martins, R. Lockwood, J. P. Swaddle, and G. Hunt. 2022. Temporal shifts in ostracode sexual dimorphism from the Late Cretaceous to the late Eocene of the US Coastal Plain. Marine Micropaleontology, 101959. Download pdf Copy.

Beane, R. J., E. M. D. Baer, R. Lockwood, R. H. Macdonald, J. R. McDaris, V. R. Morris, I. J. Villalobos, and L. D. White. 2021. Uneven increases in racial diversity of US geoscience undergraduates. Communications Earth and Environment 2: 136. Download pdf Copy. 

Martins, M. João Fernandes, G. Hunt, C. Milagros Thompson*, R. Lockwood, John P. Swaddle, and T. M. Puckett. 2020. Shifts in sexual dimorphism across a mass extinction in ostracods: implications for sexual selection as a factor in extinction risk. Proceedings of the Royal Society B 287: 20200730. Download pdf Copy.

Lockwood R, Mann R. 2019. A conservation palaeobiological perspective on Chesapeake Bay oysters. Phil. Trans. R. Soc. B 374: 20190209. Download pdf Copy.

Lukes, L. A., K. Ryker, C. Millsaps*, R. Lockwood, M. D. Uhen, C. George, C. Bentley, and P. Berquist. 2019. Leveraging a large database to increase access to undergraduate research experiences. Scholarship and Practice of Undergraduate Research, 2: 4-13. Download pdf Copy.

*Zimmt, Joshua B., R.  Lockwood, C. F. T. Andrus, and G. S. Herbert. 2019. Sclerochronological basis for growth band counting: A reliable technique for life-span determination of Crassostrea virginica from the mid-Atlantic United States. Palaeogeography, Palaeoclimatology, Palaeoecology 516: 54-63. Download pdf Copy.

Cohen, P. A., L. Park Boush, and R. Lockwood, eds. 2018. Pedagogy and Technology in the Modern Paleontology Classroom. Essentials of Paleontology. Cambridge University Press

Cohen, P.A., R. Lockwood, S. Peters. 2018. Integrating Macrostrat and Rockd into undergraduate earth science teaching. Essentials of Paleobiology Cambridge University Press. Download pdf Copy.

Lockwood, R., P. A. Cohen, M. D. Uhen, K. Ryker. 2018. Utilizing the Paleobiology Database to provide educational opportunities for undergraduates. Essentials of Paleontology. Cambridge University Press. Download pdf Copy.

Fernandes Martins, M. J., T. M. Puckett, R. Lockwood, J. P. Swaddle, G. Hunt. 2018. High male sexual investment as a driver of extinction in fossil ostracods. Nature 556: 366-369. Download pdf Copy.

Ivany, L. C., C. Pietsch, J. C. Handley, R. Lockwood, W. D. Allmon, and J. A. Sessa. 2018. Little lasting impact of the Paleocene-Eocene Thermal Maximum on shallow marine molluscan faunas. Science Advances, 4(9), eaat5528. Download pdf Copy.

*Kusnerik, K. M., R. Lockwood, A. N. Grant*. 2018. Using the fossil record to establish a baseline and recommendations for oyster mitigation in the Mid-Atlantic U.S. Marine Conservation Paleobiology. C. L. Tyler, C. L Schneider, eds. Springer–Verlag. Download pdf Copy.

Fernandes Martins, M. J., G. Hunt, R. Lockwood, J. P. Swaddle, D. J. Horne. 2017. Correlation between investment in sexual traits and valve sexual dimorphism in Cyprideis species (Ostracoda). PLOS One. 12(7), e0177791. Download pdf Copy.

Hunt, G., M. J. Fernandes Martins, T. M. Puckett, R. Lockwood, J. P. Swaddle, C. M. S. Hall*, J. Stedman. 2017. Sexual dimorphism and sexual selection in cytheroidean ostracodes from the Late Cretaceous of the US Coastal Plain. Paleobiology 43(4), 620-641. Download pdf Copy.

Canuel, E. A., G. Brush, T. M. Cronin, R. Lockwood, and A. R. Zimmerman. 2017. Paleoecology studies in Chesapeake Bay: A model system for understanding interactions between climate, anthropogenic activities, and the environment. Applications of Paleoenvironmental Techniques in Estuarine Studies. K. Weckström, K. M. Saunders, P. A. Gell, C. G. Skilbeck, eds. Developments in Paleoenvironmental Research 20: 495-527. Download pdf Copy.

Rick, T. C., L. A. Reeder-Myers, C. A. Hofman, D. Breitburg, R. Lockwood, G. Henkes, ... & M. B. Ogburn. 2016. Millennial-scale sustainability of the Chesapeake Bay Native American oyster fishery. Proceedings of the National Academy of Sciences. DOI:10.1073/pnas.1600019113. Download pdf Copy.

Carrano, M. T., M. P. Oreska*, and R. Lockwood. 2016. Vertebrate paleontology of the Cloverly Formation (Lower Cretaceous), II: Paleoecology. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2015.1071265. Download pdf Copy

*Orzechowski, E. A., R. Lockwood, J. Byrnes, S. C. Anderson†, S. Finnegan, Z. V. Finkel, P. G. Harnik, D. R. Lindberg, L. H. Liow, H. K. Lotze, C. R. McClain, J. L. McGuire‡, A. O’Dea, J. M. Pandolfi, C. Simpson‡, and D. P. Tittensor. 2015. Ecological and environmental determinants of extinction risk in the marine fossil record. Global Change Biology. DOI: 10.1111/gcb.12963. Download pdf Copy.

Finnegan, S., S. C. Anderson†, P. G. Harnik, C. Simpson‡, D. P. Tittensor, J. Byrnes, Z. V. Finkel, H. K. Lotze, D. R. Lindberg, L. H. Liow, R. Lockwood, C. R. McClain, J. L. McGuire‡, A. O’Dea, and J. M. Pandolfi. 2015. Paleontological baselines for evaluating extinction risk in the modern oceans. Science 348: 567-570. Download pdf Copy.

*McClure, K. J. and R. Lockwood. 2015. Phylogenetic relationships among Venericardia (Bivalvia: Carditidae) in North America during the Paleogene. Journal of Paleontology 89: 522-531. Download pdf Copy.

†Simonson, A. L, R. Lockwood, and J. F. Wehmiller. 2013. Three approaches to radiocarbon calibration of amino acid racemization in Mulinia lateralis from the Holocene of the Chesapeake Bay, USA.Quaternary Geochronology. Download pdf Copy

Rick, T. C. and R. Lockwood. 2013. Integrating paleobiology, archaeology, and history to inform biological conservation. Conservation Biology 27: 45-54. Download pdf Copy

‡Harnik P.G., H. K. Lotze, S. C. Anderson, Z. V. Finkel, S. Finnegan, D. R. Lindberg, L. H. Liow, R. Lockwood, C. R. McClain, J. L. McGuire, A. O’Dea, J. M. Pandolfi, C. Simpson, and D. P. Tittensor. 2012. Extinctions in ancient and modern oceans. Trends in Ecology and Evolution 27: 608-617. Download pdf Copy

Yacobucci, M. M. and R. Lockwood, editors. 2012. Teaching Paleontology in the 21st Century. Paleontological Society Special Publication 12. 256 pp

Poirier, R.K.*, Cronin, T.M., Briggs Jr., W.M., Lockwood, R., 2012, Central Arctic paleoceanography for the last 50kyr based on ostracode faunal assemblages. Marine Micropaleontology 88-89: 65-76. Download pdf Copy

‡Harnik, P. G. and R. Lockwood, 2011, Extinction in the marine Bivalvia. In Treatise on Invertebrate Paleontology Online. Number 29. Part N, Revised, Volume 1, Chapter 24. J. Carter and R. K. Thomas, eds. University of Kansas.  Download pdf Copy

*Kolbe, S. E., R. Lockwood, and G. Hunt, 2011, Does morphological variability buffer against extinction? A test using veneroid bivalves from the Plio-Pleistocene of Florida. Paleobiology 37: 355-368. Download pdf Copy

Lockwood, R. 2008. Beyond the Big Five: Extinctions as experiments in the history of life. In From Evolution to Geobiology: Research Questions Driving Paleontology at the Start of a New Century. P. E. Kelley and R. A. Bambach, eds. In press.  Download pdf Copy

Lockwood, R. and L. A. Work* 2006. Quantifying taphonomic bias in molluscan death assemblages from the upper Chesapeake Bay: patterns of shell damage. Palaios 221:442-450. Download pdf Copy

*Shelton, J., R. Lockwood, and L. M. Bybell. 2006. The effects of the Chesapeake Bay impact on calcareous nannofossil assemblages: patterns from the Watkins School Core, Newport News, Virginia (USA). Journal of Nannoplankton Research 28: 71-80. Download pdf Copy

Lockwood, R. and L. R. Chastant*. 2006. Quantifying taphonomic bias of compositional fidelity, species richness, and rank abundance in molluscan death assemblages from the upper Chesapeake Bay. Palaios 21: 376-383. Download pdf Copy

Kosnik, M. A., D. Jablonski, R. Lockwood, and P. M. Novack-Gottshall. 2006. Quantifying molluscan body size in evolutionary and ecological analyses: maximizing the return on data collection efforts. Palaios 21: 588-597. Download Pdf Copy

Lockwood, R. 2005. Body size, extinction events, and the early Cenozoic record of veneroid bivalves: a new role for recoveries. Paleobiology 31: 578-590. Download pdf Copy

Lockwood, R. 2004. The K/T event and infaunality: morphological and ecological patterns of extinction and recovery in veneroid bivalves. Paleobiology 30: 507-521. Download pdf Copy

Lockwood, R. 2003. Abundance not linked to survival across the end-Cretaceous mass extinction: patterns in North American bivalves. Proceedings of the National Academy of Sciences 100: 2478-2482. Download pdf Copy

Swaddle, J. P. and R. Lockwood. 2003. Wingtip shape and flight performance in the European Starling (Sturnus vulgaris). Ibis. 145:457-464. Download pdf Copy

Lockwood, R., J. P. Swaddle, and J. M. V. Rayner. 1998. Avian wingtip shape reconsidered: morphological adaptations to migration. Journal of Avian Biology 29: 273-292. Download pdf Copy

Swaddle, J. P. and R. Lockwood. 1998. Morphological adaptations to predation risk in passerines. Journal of Avian Biology 29: 172-176. Download pdf Copy


110: Earth's Environmental Systems

150W: Extinction is Forever

303: Age of Dinosaurs

310: Tropical Coral Reef Environments of the Bahamas

423: Paleontology

426: Evolution and the Fossil Record