Areas of Specialization
Developmental Biology: Cell movement and adhesion in embryonic development.
Evolution: Industrial melanism in American peppered moths.
Higher Education: Academic leadership, intercollegiate athletics, science education.
Rock Art: Ancient rock art representations of birds in the American Southwest.
A.B., Hiram College, 1966; M.A., Ph.D., Princeton University, 1969, 1970
Visiting Professor of Biology, Colorado State University, 2008-
Assistant, Associate, Professor of Biology, College of William & Mary, 1971, 1977,
1986-2004 (Chair, 1982-87, 1988-93, 1996-2004)
Special Assistant to the Provost, College of William & Mary, 1989
Special Assistant to the President, University of Colorado, 1987-88
Visiting Scientist, Human Leukemia Program, The Ontario Cancer Institute and The
University of Toronto, 1974-75